Geotropism of Agriolimax

On an inclined glass plate the slug Agriolimax orients and creeps upward or downward. The angle of orientation on the plane (theta) is proportional to the logarithm of the component of gravity in the creeping plane. The coefficient of variability of the measured values of (theta) decreases linearly as the logarithm at the gravity component in the creeping plane increases. The cosine of the angle of orientation decreases almost directly in proportion to the sine of the angle of inclination of the creeping plane to the horizontal, as previously found for young rats (Crozier and Pincus). But a more satisfactory formulation for the present case shows that the sine of the angle of orientation (theta) decreases in direct proportion to the increase of the reciprocal of the sine of the angle of inclination of the creeping plane. This formulation is derived from the theory that the geotropic orientation is limited by the threshold difference between the pull of the body mass on the mutually inclined longitudinal muscles at the anterior end of the slug.